Skipper, Kowalski, Rico and Private join forces with undercover organization The North Wind to stop the villainous Dr. Octavius Brine from destroying the world as we know it.

Next stop: New York! But to get back home, Alex and friends must hitch a ride with a traveling European circus and evade an evil animal-control officer.

Four animal friends get a taste of the wild life when they break out of captivity at the Central Park Zoo and wash ashore on the island of Madagascar.