America, 1890. Johnny Joestar, a genius jockey now paralyzed from the waist down, chases after outlaw Gyro Zeppeli in a tough cross-continent horse race.

Iroha's life gets knocked off its orbit when Kaguya, a carefree runaway from the Moon, moves in and convinces her to perform in a virtual world together.

After rebel forces attack Earth, a carefree heir becomes the prime suspect and joins a special military unit to find the true mastermind on the Moon.